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No one knows exactly when Gramma was born. But if the estimated birth year of 1884 is accurate, Chester Arthur occupied the Oval Office and there were only 39 states at the time.

Gramma, the San Diego Zoo’s oldest resident and one of the world’s oldest Galapagos tortoises, has died at the estimated age of 141.

The San Diego Zoo's oldest resident, a Galapagos tortoise named Gramma, died Nov. 20. She was estimated to be 141 years old.

The beloved Galapagos tortoise named Gramma that was the oldest resident of the San Diego Zoo at an estimated 141 years has died.